According to Reblitz, after touch is necessary in order for a vertical piano backchecks ( The felt cushions that hold the hammer some halfway to the strings when you play and hold a loud note ) to work properly. What happens is that some actions, when regulated to the standard dimensions of 10mm key dip and 44.5mm hammer stroke ( or distance to the strings ) won't work right that way, something about the jack not tripping far enough from where it pushes on the hammer butt.
So a little aftertouch, like 11mm key dip instead of 10mm, won't make the action feel weird and will fix these problems, although more than that does feel strange.
Now, again, that is for an UPRIGHT. Properly regulated grands aren't supposed to have ANY after touch, I believe.